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virus protein hamster position 3/rsv replication titer vero cells vaccine antibody animal genome 3/hmpv position 1 chimeric piv 3 genome table level neutralizing hmpv f protein infection expression sequence assay tract tissue metapneumoviru study rsv f challenge hpiv 3 rsv g hmpv /nl children virus replication wild-type titre chimeric viruses glycoprotein p.f.u 0.1 position 2 syncytial virus rhesus monkeys insertion antigen nl /1/00 virol plasmid fragment gene insertions result attenuated neutralizing antibody titers lysate subgroup vector plaque candidate response hmpv replication 0.5 serum rsv f protein nl /1/99 polyclonal minigenome parainfluenza ml 21 leader plaque assays wild-type rsv presence dilution 106 pfu fluenza parain hmpv f gene neutralization /1/00
