For half a century it was assumed that granule cells use ultra-sparse encoding, but now in vivo calcium-imaging studies have shown that large ensembles of granule cells provide dense signals, which themselves evolve and adapt during training.

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Journal Current Biology
Badura, A.M, & de Zeeuw, C.I. (2017). Cerebellar Granule Cells: Dense, Rich and Evolving Representations. Current Biology, 27(11), R415–R418. doi:10.1016/j.cub.2017.04.009